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EARLY EARTH and the ORIGIN OF LIFE. Major Episodes. Isotopes of carbon. Life on Earth: 3.5-4.0 billion years Prokaryotes: 3.5 billion stromatolites Oxygen: 2.7 billion Cyanobacteria Eukaryotes: 2.1 billion Multicellular Eukaryotes: 1.2 billion Animal Diversity: Cambrian period
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EARLY EARTH and the ORIGIN OF LIFE
Major Episodes Isotopes of carbon
Life on Earth: 3.5-4.0 billion years • Prokaryotes: 3.5 billion • stromatolites • Oxygen: 2.7 billion • Cyanobacteria • Eukaryotes: 2.1 billion • Multicellular Eukaryotes: 1.2 billion • Animal Diversity: Cambrian period • Land plants: 500 million
THE ORIGIN OF • LIFE • Spontaneous • generation • Biogenesis • Louis Pasteur
Miller & Urey • Primitive • Atmosphere • H2O • H2 • CH4 • NH3 • Formation of organic monomers – first step in origin of life
RNA may have been the first self replicating genetic material. • Replication & ribozymes (RNA autocatalytic) • Natural selection (genotype & phenotype)
PROTOBIONTS – aggregates of abiotically produced molecules liposome
RNA template for polypeptide formation Polypeptides act as primitive enzymes that aid replication of all RNA molecules, including competing RNAs
PROKARYOTES extremophiles eubacteria
STRUCTURE & FUNCTION • Shape: cocci (spherical), bacilli (rod), spirilla (helical) • Diplo- (2), strepto- (chain), staphylo- (cluster) • Peptidoglycan (modified sugars cross-linked by short polypeptides) in cell walls • Gram stain: Gram+ (simpler walls, thick peptidoglycan) Gram- (more complex, less peptidoglycan) • Capsule: protective layer outside wall, sticky • Pili: surface appendages
Motility: flagella, spirochetes (corkscrew,) slimy threads Taxis: movement toward or away from stimulus
ORGANIZATION & REPRODUCTION • Specialized membranes • Nucleoid region • Plasmids* • Binary fission • Transformation • Conjugation • Transduction • Endospores • Antibiotics
Nutritional Diversity: • Saprobes • Parasites • Metabolism of petroleum • Nonbiodegradable (synthetic organic compounds) Oxygen use: Obligate aerobes Facultative anaerobes Obligate anaerobes • Nitrogen Metabolism: • Nitrogen fixation • N2 NH4 • Cyanobacteria
EXTREMOPHILES • Methanogens • Extreme halophiles (bacteriorhodopsin • Extreme thermophiles
ECOLOGICALIMPACT • Decomposers • Symbiosis • Mutualism • Commensalism • Parasitism • Pathogenic • Koch’s Postulates • Exotoxins • Endotoxins • Bioremediation
Characteristics • Eukaryotic, unicellular, colonial, multicellular • Nutrition: aerobic, photoautotrophs, heterotrophs, mixotrophs • Ingestive (protozoa), photosynthetic (algae), absorptive (fungus like) • Motility: flagella, cilia, psuedopodia • Life cycles: asexual (mitosis), sexual (meiosis & syngamy), cysts • Habitat: aquatic (plankton) & moist terrestrial areas, contractile vacuoles
EVOLUTIONARY TRENDS: 1) Filamentous Cyanobacteria – specialized cells 2) Complex communities –species w/ metabolic specialties 3) Compartmentalization within cells eukaryotes ENDOSYMBIOSIS
Diplomonadida and Parabasala lack mitochondria Giardia lamblia Trichomonas vaginalis
Euglenozoa • both photosynthetic and heterotrophic flagellates Euglena - mixotrophic Trypanosoma – African sleeping sickness
Alveolata • unicellular protists with subsurface cavities (alveoli) • Dinoflagellates: red tides (deadly toxins) Gonyaulax • Ciliates: paramecium, stentor • Apicomplexans: all parasitic, plasmodium - malaria
Stramenopila • The stramenopile clade includes the water molds • and the heterokont algae Oomycota Water mold Diatoms: glass like cell walls Chrysophytes: Golden algae Phaeophytes: brown algae, Seaweeds (kelps)
SEAWEEDS • Structural and biochemical adaptations help seaweeds survive and reproduce at the ocean’s margins • Food source, thickening agents, agar • Some algae have life cycles with alternating multicellular haploid and diploid generations
Rhodophyta: Red algae lack flagella Most abundant large algae in warm coastal waters of tropical oceans
Chlorophyta • Green algae and plants evolved from a common photoautotrophic ancestor • Unicellular (chlamydomonas), colonial (volvox), filamentous (spirogyra), multicellular (ulva) • Lichens
A diversity of protists use pseudopodia for movement and feeding Rhizopoda: amoebas, amoebic dysentery
Actinopoda (Heliozoan & Radiolarians) Foraminiferans calcium carbonate walls, limestone fossils, Dover cliffs
Mycetozoa: Slime molds have structural adaptations and life cycles that enhance their ecological roles as decomposers