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This course, BIOL 3240, delves into the roles of energy and nutrients in ecosystems. Students will explore trophic levels, the classifications of organisms such as autotrophs and heterotrophs, and the significance of photosynthetically active radiation (PAR). The course covers various photosynthetic pathways, including C3, C4, and CAM processes, and examines the chemical composition and nutritional requirements of organisms. Key topics also include the feeding strategies of herbivores, carnivores, and detritivores, emphasizing the interplay between energy limitation and food density in ecological interactions.
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BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Energy Sources • Organisms can be classified by trophic levels. • Autotrophs use… • ______________: Use CO2 as carbon source, and sunlight as energy. • ______________: Use inorganic molecules as source of carbon and energy. • Heterotrophs use…
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Solar - Powered Biosphere • Light propagates through space… • _________: Particle of light bears energy. • ____________(IR) Long-wavelength, low energy. • ____________(UV) Short wavelength, high energy. • _____________________________________(PAR)
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Photosynthetically Active Radiation
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Solar - Powered Biosphere • PAR • Quantified as _____________________. • Number of photons striking square meter surface each second. • Chlorophyll absorbs light as photons. • Landscapes, water, and organisms can all change the amount and quality of light reaching an area.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Photosynthetic Pathways • Used by… • CO2+ ribulosebisphosphate (5 carbon sugar) = phosphoglyceric acid (3 carbon acid)
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients C3 Photosynthesis
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Photosynthetic Pathways • Reduce internal CO2 concentrations. • Acids produced during carbon fixation diffuse to specialized cells surrounding _______________.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients C4 Photosynthesis
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Photosynthetic Pathways • (Crassulacean Acid Metabolism) • Limited to succulent plants in arid and semi-arid environments.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients CAM Photosynthesis
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Using Organic Molecules • Three Feeding Methods of Heterotrophs: • _______________: Feed on plants. • _______________: Feed on animal flesh. • _______________: Feed on non-living organic matter.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Chemical Composition and Nutrient Requirements • Five elements make up 93-97% of biomass of plants, animals, fungi and bacteria:
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Essential Plant Nutrients • Potassium • Calcium • Magnesium • Sulfur • Chlorine • Iron • Manganese • Boron • Zinc • Copper • Molybdenum
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Herbivores • Substantial nutritional chemistry problems. • Low ... • Must overcome plant physical and chemical defenses. • Physical • Chemical
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Detritivores • Consume food rich in ___________________, but poor in ______________. • Fresh detritus may still have considerable ________________________ present.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Carnivores • Consume … • Cannot choose prey at will. • Prey Defenses:
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Carnivores • Predators are usually selection agents for refined prey defense. • Usually eliminate more conspicuous members of a population (less adaptive). • Must catch and subdue prey - ________________. • Predator and prey species are engaged in a co-evolutionary race.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Using Inorganic Molecules • 1977 - Organisms found living on sea floor. • Near nutrients discharged from volcanic activity through oceanic rift. • Autotrophs depend on chemosynthetic bacteria.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Energy Limitation • Limits on potential rate of energy intake by animals have been demonstrated by studying relationship between feeding rate and food availability. • Limits on potential rate of energy intake by plants have been demonstrated by studying response of photosynthetic rate to photon flux density.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Photon Flux and Photosynthetic Response Curves • Rate of photosynthesis increases linearly with photon flux density at … , rises more slowly with __________________ , and tends to level off at _________________. • Response curves for different species generally level off at _______________ maximum photosynthesis rates.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Food Density and Animal Functional Response • Holling described (3) basic functional responses: • 1. Feeding rate increases linearly as food density increases - levels off at maximum. • 2. Feeding rate rises in proportion to food density.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Food Density and Animal Functional Response • 3. Feeding rate increases most rapidly at intermediate densities
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Optimal Foraging Theory • Assures if energy supplies are limited, organisms cannot simultaneously maximize all life functions. • Must compromise between competing demands for resources.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Optimal Foraging in Bluegill Sunfish
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Optimal Foraging Theory • All other things being equal,more abundant prey yields larger energy return. Must consider energy expended during: • Tend to maximize rate of energy intake.
BIOL 3240 Plant and Animal Ecology – Energy & Nutrients Optimal Foraging By Plants • Limited supplies of energy for allocation to … • Bloom suggested plants adjust allocation in such a manner that all resources are equally limited. • Appear to allocate growth in a manner that increases rate of acquisition of resources in ______________________.