Theme 2 - Implementation

1. Theme 2 - Implementation Theme 2 has 34 Tasks 49 Deliverables Theme 2: Ecosystem impacts, acclimation and adaptation CO2 Climate relevant gases WP5 WP8Trophic interactions Ocean chemistry Autotrophic processes WP7 Microbial activity WP4-8 Adaptation WP4 Biocalcification Nutrient availability WP6 Reproduction & growth WP9 From process studies to ecosystem models

2. WP4 Sensitivity of calcification Theme 2 - Implementation Organism responses Community level responses lab experiments Svalbard experiment WP5 Sensitivity of autotrophic processes WP7 Microbial diversity and activity, in particular nitrogen cycling WP8 Impact on trophic interactions WP6 Performance: reproduction and growth WP9 From process studies to ecosystem models

3. Theme 2 - Implementation Organism responses Community level responses WP4 Sensitivity of calcification Calcification mechanisms in coccolithophores, foraminifera, pteropods, corals, and bivalves Physiological responses of marine calcifiers to ocean acidification at the species and community levels using genomic and proteomic approaches Rates of genetic change in calcifying genes in key calcifiers from decadal to geological time scales Effect of the phytoplankton community composition on the ballast effect

4. Theme 2 - Implementation Organism responses Community level responses microcosm experiments lab experiments Svalbard experiment WP5 Sensitivity of autotrophic processes WP7 Microbial diversity and activity, in particular nitrogen cycling WP8 Impact on trophic interactions WP5 C fixation, exudation, POC/DOC, stoichiometry, CR biogases, bacterial gene expr. WP5 Microalgal C fluxes and engery allocation – cell model WP7 prokaryote-mediated carbon flow prokaryotic diversity and gene expression WP7 bioturbation activity, its influence on microbial diversity and nitrogen recycling WP8predator-mediated mortality, effects of prey quality algal exudates  aggregation control by grazing & viral lysis role of coccoliths in reducing grazing and viral mortality WP5 speciation and bioavailability of major and trace nutrients

5. Theme 2 - Implementation Organism responses Community level responses WP6 Performance: reproduction and growth Experiments on key organisms: copepods, bivalves, bioturbators, squid & fish Sensitivity of early life stages Acid-base regulation  changes in gene expression CO2 effects on temp. tolerance  latitudinal cline adaptation potential

6. Theme 2 - Implementation Organism responses Community level responses WP9 From process studies to ecosystem models Process oriented meta-analysis of experimental results Modeling of mesocosm experiments, including data-assimilative parameter estimation Meta-analysis of model structures and parameterizations Models improvement for optimal balance between model complexity, performance and portability Parameterization assessment  recommendations for incorporation into global-scale models

7. Svalbard experiment National representatives: Germany Ulf Riebesell/Hans Pörtner France Markus Weinbauer UK Steve Widdicombe Norway Richard Bellerby/Frede Thingstad Netherlands Corina Brussaard S. Korea Kitack Lee Italy ? China ? Find national funding to supplement funding for Svalbard experiment, e.g. to cover bench fees, transportation, …