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Discover the characteristics, classes, feeding, reproduction, and systems of Annelids - Polychaeta, Oligochaeta, and Hirudinea - segmented worms with significant ecological roles.
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Today’s Objectives • Explore the diversity, success, body plans, organ systems and economic importance of phylum Annelida.
Annelida Segmented Worms
Main Characteristics • Bilateral symmetry • Metameric • Tagmatization • Protostomes • Triploblastic • Setae • Closed Circulatory System • Ganglia and Nerve Cords • Metanephridia
Classes of Annelids • Polychaeta – Nereis, Arenicola • Oligochaeta – Lumbricus, Tubifex • Hirudinea - Hirudo
Class Polychaeta • Mostly marine • Parapodia • Cuticle • 1st segment – prostomium • 2nd segment – peristomium
Polychaeta Feeding • Most carnivorous • Some have venom • Gut is straight tube • Some are detritovores • Extract nutrition from sediment (or soil) • Gut has crop and gizzard • Many are filter feeders • Some can get food by diffusion
Other polychaete Systems • Respiration by diffusion • 2-4 pairs of eyes on/near prostomium • Chemoreceptors • Statocysts • Metanephridium for excretion • Chloragogen tissue for protein metabolism
Polychaete Reproduction • Can regenerate • Asexual reproduction • Budding • Fission • Sexual reproduction • Most fertilization external • Can have alternation of generations • Epitoke (sexual) vs. Atoke (asexual)
Oligochaeta • Terrestrial, freshwater, some marine • Have a clitellum • For mucus secretion • Used in copulation • Used to form cocoons • No parapodia • Few setae
Oligochaete Locomotion • Use circular and longitudinal muscles • Can use setae as anchors • Use hydrostatic pressure
Oligochaete Feeding • Scavengers and/or detritovores • Path of food • Mouth • Pharynx • Crop • Gizzard • Stomach • Intestine
Other Oligochaete Systems • Most ganglia fused • Reduced eyes • Sensitive to chemical or mechanical stimuli • Use metanephridia for excretion • Chloragogen tissue
Oligochaete Reproduction • Hermaphroditic • Must line up clitella • Held together by mucus sheath • Cocoon formed • Fertilization occurs here • No larval stages • Some freshwater species asexual
Hirudinea • Terrestrial, freshwater or marine • No parapodia • Secondary annuli on segments • Circular, longitudinal and oblique muscle layers
Hirudinean Feeding • Many carnivorous • Small invertebrates • Body fluids • Mouth in the anterior sucker • Produce “hirudin” – anticoagulant
Other Hirudinea Systems • Gas exchange by diffusion • Nervous system • Photoreceptors • Can sense temperature • Sensory papillae • 10-17 pairs of metanephridia for waste • Chloragogen tissue
Hirudinea Reproduction • All monecious • All sexually reproducing • Have a penis for sperm transfer • Clitellum seen during breeding season • No larval stages