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Outline. Energy flow versus chemical cycles Carbon Nitrogen Phosphorous Water Ecosystem services. Energy flow through ecosystems. Only 5% of the solar energy that arrives on earth is captured by photosynthesis. The energy captured powers life. Trophic levels. Energy flow versus
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Outline • Energy flow versus chemical cycles • Carbon • Nitrogen • Phosphorous • Water • Ecosystem services
Energy flow through ecosystems Only 5% of the solar energy that arrives on earth is captured by photosynthesis. The energy captured powers life.
Energy flow versus Biogeochemical cycles
A generalized biogeochemical cycle Consumers Producers Decomposers Abiotic reservoir Geological processes
Carbon cycle Carbon is the currency for energy flow!
Carbon cycle Carbon is the currency for energy flow! Carbon cycles through the atmosphere, organisms, soils, and oceans (and rocks at very long time scales)
Carbon cycle Carbon is the conduit for energy flow! Carbon cycles through the atmosphere, organisms, soils, and oceans (and rocks at very long time scales) Gaseous, mineral, organic
CO2, C leaching, erosion NPP Soil, plant growth (wood) Ecosystem carbon balance C in = storage + C out C in > storage + C out: accumulating C C in = storage + C out: balanced C C in < storage + C out: loosing C
Carbon stocks Sedimentary rocks > deep oceans > atmosphere Only a small amount in organisms and soils!
Nitrogen cycle • N cycles through the atmosphere, organisms, soils and sediments
Nitrogen cycle • N cycles through the atmosphere, organisms, soils and sediments • Organically bound N is one of the main constituents of organisms • enzymes for photosynthesis, metabolism • proteins • building blocks of DNA...
Nitrogen cycle • N cycles through the atmosphere, organisms, soils and sediments • Organically bound N is one of the main constituents of organisms • enzymes for photosynthesis, metabolism • proteins • building blocks of DNA... • Gaseous, mineral, organic
Nitrogen cycle • Although N2 gas makes up 72% of the atmosphere, few organisms can use it. • N is the element that is most likely to limit plant growth
Nitrogen cycle • N enters system through biological fixation
Picture of N accumulation over primary succession Nitrogen accumulates over primary succession
Nitrogen cycle • N enters system through biological fixation • If organisms can fix N, why don’t they take over and fix enough N so that it is no longer limiting to plant growth?
Why is N in short supply? • Only a few types of organisms can do it
Why is N in short supply? • Only a few types of organisms can do it • Energetically expensive to fix N
Why is N in short supply? • Only a few types of organisms can do it • Energetically expensive to fix N • Chemically expensive to fix N
Why is N in short supply? • Only a few types of organisms can do it • Energetically expensive to fix N • Chemically expensive to fix N • N can be lost from the system • Leaching • Fire • Erosion
Hydrological cycle Water cycles through the oceans, fresh waters, atmosphere, and land
Hydrological cycle Water cycles through the oceans, fresh waters, atmosphere, and land Gaseous phase and liquid phase
Hydrological cycle Evaporation: liquid to gas Transpiration: plant transport of water from the soil to the atmosphere Precipitation: gas to liquid
Water inputs to ecosystems • Precipitation • THE major water input to ecosystems • Groundwater (in some ecosystems) • Fog deposition (in some ecosystems)
Water balance in ecosystems Inputs = storage + outputs
Water balance in ecosystems Soil is like a bucket
Water balance in ecosystems Soil is like a bucket • Inputs • Mainly precipitation
Water balance in ecosystems Soil is like a bucket • Inputs • Mainly precipitation • Storage • Plants, soil and water bodies
Water balance in ecosystems Soil is like a bucket • Inputs • Mainly precipitation • Storage • Plants, soil and water bodies • Outputs • Evaporation • Transpiration • Drainage
Hydrological cycle • Over oceans, evaporation >> precipitation • On land, evaporation << precipitation and transpiration • Lakes, rivers, streams, groundwater
Ecosystem services Processes by which the environment produces fundamental life-support services enjoyed by humans
Ecosystem services • Purification of air and water
Ecosystem services • Purification of air and water • Detoxification and decomposition of wastes
Ecosystem services • Purification of air and water • Detoxification and decomposition of wastes • Regulation of climate
Ecosystem services • Purification of air and water • Detoxification and decomposition of wastes • Regulation of climate • Regeneration of soil fertility
Ecosystem services • Purification of air and water • Detoxification and decomposition of wastes • Regulation of climate • Regeneration of soil fertility • Production and maintenance of biodiversity
Ecosystem services Key ingredients of our agricultural, pharmaceutical, and industrial enterprises
Ecosystem services We know that they are: • Essential to civilization
Ecosystem services We know that they are: • Essential to civilization • So complex that they can’t be replaced by technology (???)
Ecosystem services We know that they are: • Essential to civilization • So complex that they can’t be replaced by technology (???) • Impaired by human activities