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Chapter 25 Phylogeny and the Tree of Life. Chapter focus. Shifting from the process of how evolution works to the pattern evolution produces over time. Phylogeny. Phylon = tribe, geny = genesis or origin
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Chapter focus • Shifting from the process of how evolution works to the pattern evolution produces over time.
Phylogeny • Phylon = tribe, geny = genesis or origin • The evolutionary history of a species or a group of related species.
Systematics • The study of biological diversity and classification. • Uses evidence from the fossil record and other sources to reconstruct phylogeny.
Systematics fuses: 1. Phylogeny- tracing of evolutionary relationships. 2. Taxonomy- the identification and classification of species.
Taxonomy • Natural to humans. • Modern system developed by Linnaeus in the 18th century.
Scientific names • Composed of Genus and species. • Written in Latin and shown in italics or underlined. • Governed by a set of rules and procedures.
Linnaeus Taxonomy 1. Binomial Nomenclature – two names for each organism. Ex - Homo sapiens 2. Hierarchical System – arranges life into groups. Ex - Kingdom Species
Goal of Systematics • To have Taxonomy reflect the evolutionary affinities or phylogeny of the organisms.
Phylogenetic Tree • Branching diagram showing evolutionary relationships between organisms.
Trees show: • Ancestral lineage • Branch points or nodes • Length of branch point suggests “time” and degree of closeness.
Phylogenetic tree - Example Branch point Ancestral Lineage
Question? • How to group taxa so that the phylogenetic relationships are correct ?
Ideal Situation • Monophyletic Grouping - a single ancestor gave rise to all species in the taxon.
Other Possibilities • Polyphyletic - grouping where members are derived from two or more ancestral forms. • Paraphyletic - grouping that does not include all members from an ancestral form.
Problem • Not all “likeness” is inherited from a common ancestor. • Problem is of homology vs analogy.
Homology and Analogy • Homology – likeness attributed to shared ancestry. • Ex: forelimbs of vertebrates • Analogy – likeness due to evolution solution for the same problem. • Ex: wings of insects and birds
Convergent Evolution • When unrelated species have similar adaptations to a common environment. A specific example of Analogy. • Ex: Sharks and dolphins
Need • Methods to group organisms by similarities and phylogenies. • One possible method is Molecular Systematics.
Molecular Systematics • Compares similarities at the molecular level. • Ex: DNA, Proteins
DNA patterns • If similar DNA – more closely related, more recent common ancestor. • If different DNA – less closely related, less recent common ancestor.
Making a Phylogenetic Tree • May use morphology, genetic and other data. • Typically “rooted” in a common ancestor. • Uses statistical analysis looking for “best fit”.
Best Fit • Maximum parsimony – requires fewest DNA base changes. • Branch lengths – suggest the closeness of the relationships and the time of branch points.
Evolutionary History • Is in the organism’s genome. • Note – taxonomic relationships can be changed based on what the DNA tells us.
Result • Taxonomy will become Genealogies, reflecting the organism’s "Descent with Modification“.
Kingdom • Highest Taxonomic category • Old system - 2 Kingdoms 1. Plant 2. Animal
5 Kingdom System • R.H. Whittaker - 1969 • System most widely used, but is changing.
Main Characteristics • Cell Type • Structure • Nutrition Mode • Problems in Kingdom Monera and Protista
Current Views • Multiple Kingdoms – split life into as many as 8 kingdoms. • Domains – a system of classification that is higher than kingdom.
3 Domain System • Based on molecular structure for evolutionary relationships. • Prokaryotes are not all alike and should be recognized as two groups.
3 Domains 1. Bacteria – prokaryotic. 2. Archaea – prokaryotic, but biochemically similar to eukaryotic cells. 3. Eucarya – the traditional eukaryotic cells.
Summary • What is Phylogeny? • What is a phylogenetic tree? • How is molecular data used in phylogeny? • Know the Domains and the Tree of Life.